By D. Dudley Williams (auth.), John F. Flannagan, K. Eric Marshall (eds.)
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Extra resources for Advances in Ephemeroptera Biology
Labium: submentum (postmentum) (P) with thick lateral setae (Fig. 28). (0) lateral margins bare (Fig. 29). Body of nymph (0) robust. (P) flattened. anteromedian emargination subapical pectinate setae 47 PHYLOGENETIC RELATIONSHIPS OF LEPTOPHLEBIIDAE Table 2. Phyletic lineages representing the cool-adapted Leptophlebiidae of Southern South America and selected related genera from other areas of the Southern Hemisphere. Lineage l. HapsiphZebia Genera Included Continent HapsiphZebia AtaZophZebia, AtaZomicria Jappa, UZmerophZebia "ZephZebia" cruentata Aprionyx New Zealand Africa South America Australia " 2.
Figure 24, maxilla of New genus A. 50 MANUEL L. PESCADOR AND WILLIAM L. PETERS sister group retained the primitive states (Table 1). Progressive marginal curvature of the labrum in the five lineages forms a phenocline ranging from subparallel in the HapsiphZebia lineage (Fig. 15) through rounded in the PenaphZebia (Fig. 16) and AtaZoneZZa (Fig. 17) lineages, to angular in the Genus A (Fig. 19) and MePidiaZaPis (Figs. 20 and 22) lineages, suggesting that the character state in the MeridiaZaPis is highly derived.
These nymphs could be of great importance in the study of mayfly evolution, and therefore, an understanding of their systematic position is crucial in evaluating the data which can be obtained from their study. In a critical review of these nymphs, Demoulin (1970) removed them from the Ephemeroptera and considered them to belong in the order Archodonata. His argument that they are not Ephemeroptera depended primarily on the presence of segmented tarsi and double tarsal claws, and details of the wing venation, in particular a simple MA, separate R4 and R5, and the existence of a connecting vein between MP and CuA.